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Cover Illustration: Autofluorescence of tetrads and elaters of the hornwort Leiosporoceros dussii (Steph.) Hässel viewed with a Leica CTR5000 fluorescence microscope. The smooth bean-shaped spores are in bilateral-alterno opposite tetrads (yellowgreen) and are interspersed with long elaters (yellow-green); both spores and elaters contain plastids (red). Leiosporoceros has an assortment of unique features including spore shape and arrangement and Nostoc strand development and ultrastructure, that support its sister relationship to all other hornworts. See Villarreal and Renzaglia: Structure and development of Nostoc strands in Leiosporoceros dussii (Anthocerotophyta): a novel symbiosis in land plants, 693-705. Image taken by J. C. Villarreal A. and designed by Andrew Blackwell.