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Cover Illustration: Jubelina rosea (Malpighiaceae) from French Guiana. Malpighiaceae have been difficult to place phylogenetically owing in part to their highly specialized floral morphology, a likely adaptation to their association with oil-bee pollinators. See Davis and Chase: Elatinaceae are sister to Malpighiaceae; Peridiscaceae belong to Saxifra gales, pp. 262-273 in this issue. Photo credit: Carol Gracie.