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Cover Illustration: A phylogeny of angiosperms based on matK, a plastid gene nested within the trnK intron. Illus~ trations are superimposed on photographs of representative taxa from major angiosperm lineages, with Amborella shown in the center. We thank Peter K. Endress for providing photographs of Chloranthus and Ceratophyllum, Porter P. Lowry II for Amborella, and Duncan M. Porter for Tiarella (Saxifragaceae).